Protomantises were an advanced class of Nefarious troopers, serving as the minions of Dr. Nefarious and Lord Veroselon. It had the same "brain" like Protoguards, Splitterbots, Arc Strikers, and Cyclobombers, but was a different shape, had two small near-unnoticeable eyes, and had four small legs.
The Protomantises attacked Ratchet by hitting him with a continuous stream of fire from its twin mini-guns that were connected to its head. They would also melee him if he came too close. If hit with a Mag-Net they would sometimes still fire their twin miniguns.
After Ratchet arrived on Vorselon's Warship in order to save Alister Azimuth, the robotic troopers were replaced with stronger Chronoguard variants, taking more hits to destroy them and had blue armor instead of red. A Protomantis was first encountered on Zolar Forest, Planet Quantos. Unlike the other robots, the Protomantis never talks.